Fronds deeply tripinnatifid or tripinnate, evergreen; first pinnule (nearest the rachis of the frond) on the lower side of each basal pinna shorter than the second pinnule on the same side; indusia, rachis of frond, and rachis of pinnae with minute, stalked glands.

Fronds less finely divided, not evergreen; first pinnule on the lower side of each basal pinna longer than the second pinnule on the same side; glands absent.